Three scientists who helped establish the field of ethology (and who all shared the Nobel Prize in 1973) were Dutch ethologist (scientist who studies animals’ behavior in their natural habitat) and ornithologist Niko Tinbergen (1907–1988), Austrian ethologist and zoologist Konrad Lorenz (1903–1989), and Austrian zoologist, entomologist, and ethnologist Karl von Frisch (1886–1982)—all who lay the four cornerstones of ethological study: causation, development, evolution, and function of behavior. Lorenz studied imprinting behavior of birds, von Frisch worked on the “dance” of the honeybees, and Tin-bergen studied aggressive behavior of stickleback fish.