Thomas Huxley (1825–1895) was a staunch supporter of Darwin’s work; in fact, Huxley wrote a favorable review of Darwin’s On the Origin of Species that appeared soon after its publication. When the firestorm of controversy began after the appearance of Darwin’s book, Huxley was ready and able to defend Darwin, whose chronic public reticence about his theories was at that time exacerbated by illness. Huxley’s defense of Darwin was so vigorous during a debate with Bishop Samuel Wilberforce (1805–1873) at the British Association for the Advancement of Science in 1860 that he earned the title “Darwin’s bulldog.”